Scottish biologist D'Arcy Thompson pioneered the study of growth patterns in both plants and animals, showing that simple equations could explain spiral growth. The German biologist and artist Ernst Haeckel painted hundreds of marine organisms to emphasise their symmetry. In the 19th century, the Belgian physicist Joseph Plateau examined soap films, leading him to formulate the concept of a minimal surface. The modern understanding of visible patterns developed gradually over time. Early Greek philosophers studied pattern, with Plato, Pythagoras and Empedocles attempting to explain order in nature. Natural patterns include symmetries, trees, spirals, meanders, waves, foams, tessellations, cracks and stripes. These patterns recur in different contexts and can sometimes be modelled mathematically. Patterns in nature are visible regularities of form found in the natural world. Patterns of the veiled chameleon, Chamaeleo calyptratus, provide camouflage and signal mood as well as breeding condition.
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